Examines the influence of biological and physical processes on elemental cycling and geochemical transformations from the molecular to the global scale, involving microorganisms, higher plants and animals and whole ecosystems; factors that regulate element cycles including soils, climate, disturbance and human activities; interactions among the biosphere, hydrosphere, lithosphere, and atmosphere; transformations of C, N, S, and trace elements. Prereq: one semester biology and two semesters of chemistry or permission.